Tom McGrath

Character: Skipper (voice)

Chris Miller

Character: Kowalski (voice)

Christopher Knights

Character: Private (voice)

Conrad Vernon

Character: Rico (voice)

John Malkovich

Character: Dave (voice)

Benedict Cumberbatch

Character: Classified (voice)

Ken Jeong

Character: Short Fuse (voice)

Annet Mahendru

Character: Eva (voice)

Peter Stormare

Character: Corporal (voice)

Andy Richter

Character: Mort (voice)

Danny Jacobs

Character: King Julien (voice)

Werner Herzog

Character: Documentary Filmmaker (voice)

Sean Charmatz

Character: Cricket (voice)

Billy Eichner

Character: New York Reporter (voice)

Stephen Kearin

Character: Pilot / Aquarium Employee (voice)

Ava Acres

Character: Girl at Zoo (voice)

Kelly Cooney Cilella

Character: Mermaid Penguin (voice)

Susan Fitzer

Character: Antarctic Penguin (voice)

Chris Sanders

Character: Antarctic Penguin (voice)

Emily Nordwind

Character: Antarctic Penguin (voice)

Mike Mitchell

Character: Antarctic Penguin (voice)

Hope Levy

Character: Antarctic Penguin / Woman at Aquarium (voice)

Walt Dohrn

Character: Antarctic Penguin (voice)

Jim Pirri

Character: Gondolier (voice)

Elizabeth Pan

Character: Shanghai TV Reporter (voice)

Jeff Fischer

Character: Security System Voice (voice)

Stephen Apostolina

Character: Penguin Prisoner (voice)

Al Rodrigo

Character: Penguin Fan (voice)

Steve Alterman

Character: Aquarium Announcer (voice)

Carter Hastings

Character: Boy at Aquarium (voice)

Cullen McCarthy

Character: Boy at Zoo (voice)

Nicholas Guest

Character: Flight Attendant (voice)

Adriano Aragon

Character: Angry Man (voice)

Lynnanne Zager

Character: Computer Voice (voice)

Angie Wu

Character: Girl with Snow Globe (voice)